The original line-up consisted of: Stella Anscombe (vocals), Leoni Nicol (vocals), John Ellis (vocals), Paul Hayward (Guitar), Mark Gresty (bass) and Wayne Calcutt (drums),

After recording a first BBC session for the John Peel show (broadcast October 4, 1978), Nicol and Anscombe left the band and were replaced by Carole Brooks and Tracy Spencer.

Following a second BBC performance recorded January 31, 1979, The Molesters were signed to East London Independant label Small Wonder Records in early 1979, where they released two singles. Discouraged by tepid reviews and modest sales figures, the band dwindled to a three-piece by May 1979, consisting of Ellis, Gresty and Calcutt.

During the second half of 1979, The Molesters continued with a reformed line-up consisting of Tom Maltby (lead vocals), Paul Hayward (guitar/backing vocals), Wayne Calcutt (drums) and Paul Martin (bass), before finally calling it a day by the end of that year.

Mark Gresty and Paul Martin went on to form The Relatives who later became Life Size Models.